Zuul, Destroyer of Shins
Royal Ontario Museum
Reviewed: 9/5/2026
Museum overview of the exceptionally preserved Zuul crurivastator holotype held at the ROM, covering its anatomy, armour, tail club and discovery.
Open sourceZuul crurivastator
Zuul crurivastator was a heavily armoured, barrel-bodied plant-eater that walked on four short, pillar-like legs. It measured roughly six metres from snout to tail tip and carried rows of bony plates, called osteoderms, embedded in the skin of its back and neck.
Zuul
Zuul crurivastator
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Quick facts
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Weight
1800–2500 kg
Length
5.5–6.5 m
Diet
Herbivore
Period
Cretaceous
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1800–2500 kg
5.5–6.5 m
Cretaceous
Quadrupedal
1.2–1.5 m
10 km/h
Herbivore
One recorded fossil find, in United States. First described in 2017 by Arbour and Evans.
Sites and counts from the Paleobiology Database
The timeline shows the whole age of dinosaurs – and how much time has passed since. The glowing part is when Zuul lived.
million years ago
Lived 76.2–75.2 million years ago
If the whole age of dinosaurs were a single day, Zuul lived at 22:45.
Chapter • Meet the dinosaur
Zuul crurivastator was a heavily armoured, barrel-bodied plant-eater that walked on four short, pillar-like legs. It measured roughly six metres from snout to tail tip and carried rows of bony plates, called osteoderms, embedded in the skin of its back and neck. Blade-like spikes projected from its flanks, and the broad skull bore short horns sweeping back over the neck plus a pair of pyramid-shaped horns beneath the eyes. The whole head was sheathed in a mosaic of fused bony tiles, and the tail ended in a massive club supported by a stiffened handle of interlocking vertebrae. Like other ankylosaurids, Zuul was a slow, low-slung animal that could not outrun a predator. Its defence lay in armour and in that tail club, which could be swung sideways with considerable force. The species name crurivastator, "destroyer of shins", refers to the idea that such a blow could shatter the leg bones of an attacking tyrannosaur such as Gorgosaurus. A 2022 study of the holotype found that several spikes along the animal's flanks had been broken and had healed, while the tail club itself was undamaged. The researchers interpret this as evidence that Zuul individuals fought one another side-on, rather than that the injuries came from predators. Zuul fed on low-growing vegetation. Its broad, shovel-shaped muzzle and small, leaf-shaped teeth were suited to cropping ferns, horsetails, cycads and early flowering plants close to the ground. Ankylosaurids had a hugely capacious body cavity housing a long gut, where tough plant material could ferment and be broken down by microbes, because the teeth were not built for efficient grinding. The fossil was found in 2014 in the Judith River Formation near Havre, Montana, USA, by a crew that was actually excavating a Gorgosaurus. The skeleton – holotype ROM 75860, now at the Royal Ontario Museum in Toronto – includes a complete skull, the entire tail with its club, and much of the body with osteoderms and skin impressions preserved in life position. That makes Zuul one of the best-preserved ankylosaurs ever described. Victoria Arbour and David Evans named the species in 2017, borrowing the genus name from the monster Zuul in the 1984 film Ghostbusters because the skull reminded them of the creature's face.
Chapter • Where it lived
Zuul lived about 76 million years ago, during the Campanian age of the Late Cretaceous, on the humid coastal lowlands west of the Western Interior Seaway that then split North America in two. All known fossils come from the Judith River Formation of Montana, USA – a landscape of meandering rivers, floodplains, lagoons and forests of cycads, ferns, conifers and early flowering plants. Hadrosaurs, ceratopsians and predators such as Gorgosaurus shared the same environment.
Chapter • Wild behavior
There is no evidence that Zuul lived in herds; ankylosaurs are almost always found as isolated individuals, which is usually read as a largely solitary lifestyle. However, the healed injuries to the spikes along the holotype's flanks suggest that individuals did clash with one another, perhaps in contests over territory or mates. How common such fights were, and whether they involved only mature animals, cannot be determined from a single skeleton.
The genus is named after Zuul the Gatekeeper from the 1984 film Ghostbusters, because the short snout and backswept horns resembled the movie monster.
The species name crurivastator means "destroyer of shins", a nod to the tail club.
The holotype preserves skin, keratin sheaths and bony plates in their original positions – exceptionally rare for an ankylosaur.
The bony knob at the end of the tail was about 30 centimetres wide and sat on a stiff handle of fused vertebrae.
The specimen was discovered by chance in 2014 while a team was digging up a Gorgosaurus skeleton in Montana.
Healed damage to the flank spikes points to combat between members of the same species rather than predator attacks.
Zuul weighed roughly as much as an adult rhinoceros, around 2.5 tonnes.
The body block was nicknamed "the Zuul mummy" in the press because of its superb soft-tissue impressions.
Chapter • Scientific place
Royal Ontario Museum
Reviewed: 9/5/2026
Museum overview of the exceptionally preserved Zuul crurivastator holotype held at the ROM, covering its anatomy, armour, tail club and discovery.
Open sourcewww.ncbi.nlm.nih.gov
Reviewed: 9/5/2026
VERIFY:period_start_mya: stored=76.7 suggested=76.2 confidence=high — The holotype ROM 75860 comes from the Coal Ridge Member of the Judith River Formation, which the describing paper and subsequent 40Ar/39Ar dating (Rogers et al. 2016) place between 76.2 and 75.2 Ma. A start of 76.7 Ma predates the base of the member and is not supported.
Open sourceRoyal Society Open Science
Reviewed: 9/5/2026
Original 2017 description by Arbour & Evans erecting the genus and species and placing Zuul crurivastator phylogenetically within Ankylosauridae: Ankylosaurinae.
Open sourceRoyal Society Open Science
Reviewed: 9/5/2026
Documents the holotype locality (ROM 75860) and its stratigraphic position in the Coal Ridge Member of the Judith River Formation, Montana, USA, the only known occurrence of the species.
Open sourceThe Journal of Geology (University of Chicago Press)
Reviewed: 9/5/2026
Documents the alluvial, coastal-plain depositional environments of the Judith River Formation west of the Western Interior Seaway, the palaeoenvironment in which Zuul lived.
Open sourceRoyal Society Open Science
Reviewed: 9/5/2026
Describes the skull, jaws and dentition of Zuul crurivastator, an ankylosaurine herbivore that fed on low-growing Cretaceous vegetation.
Open sourceThe Journal of Geology (University of Chicago Press)
Reviewed: 9/5/2026
Extinct taxon; conservation status NE
Open sourceRoyal Ontario Museum
Reviewed: 9/5/2026
Museum overview of the exceptionally preserved Zuul crurivastator holotype held at the ROM, covering its anatomy, armour, tail club and discovery.
Open sourceRoyal Ontario Museum
Reviewed: 9/5/2026
Museum overview of the exceptionally preserved Zuul crurivastator holotype held at the ROM, covering its anatomy, armour, tail club and discovery.
Open sourceRoyal Ontario Museum
Reviewed: 9/5/2026
Museum overview of the exceptionally preserved Zuul crurivastator holotype held at the ROM, covering its anatomy, armour, tail club and discovery.
Open source